Well made.
Did a good job hiding the shells and hearts, I found all the hearts but I still can't find 2 shells, one of them is in Snail town (which by the way seems way to littered with secret passages for my taste), the other is in Lux Lirata (Black section of the map).
The terrain is very colorful and creative but a bit random, for instance, in Lux Lirata one minute your in the sky where there are clouds and the next your in space... in a what appears to be a space station.
Even though it is generic, I find that the way the bosses die is just awesome for NES style.
I feel like your put some thought and effort into positioning the enemies to increase the challenge, makes it hard to just whale through it all, at least until you get the better upgrades - in particular, the devastator.
In the boss rush, I don't see how the Space Box (Time Cube in boss rush) is much harder though he just seems a little faster but I'm not seeing any increase in a challenge, in fact it actually seemed a little easier to kill him. You could have given him higher endurance. The first two actually had features that made them harder in a way, the shell breaker (Super Shell Breaker in boss rush) had a ton of those circles acting like a shield against your attack, which sorta made it so you could only do serious damage when it was shooting at you because the circles disappear when it shoots.
For Stompy (Vise Vires in boss rush), not only are there more cannons shooting at you but the boss is also shooting you with those colorful ring blasts.
As for the Moon Snail (Sun Snail in boss rush) I'm not seeing any big difference in his first stage - he might be moving faster but I'm not sure, as for his second stage the changes I'm seeing aren't making him any harder to fight. But since he was a big challenge to begin with and your fighting him with only 5 hearts and a few less upgrades, I can see why you did little to change the fight against him.
The "super secret boomerang" - What so special about it? does it like super amplify the power of you boomerang or something? Also the way back out from the area where it is at, you need to kinda have to glitch jump. You can get out but you can't just slide along the wall and expect to slip through a secret passage
The music at the Shrine of Isis kinda sound like music you might find in a really old pokemon game (A "mysterious" area or something)
The main path to the are where the shrine is hidden (which by the way looks like a hacked NES level - what's up with that?) wasn't hard to find, but you could easily miss the way to actual shrine itself (I suppose THAT is the only real reason the area looks so weird then).